Humans are thought to be responsible for a large number of environmental problems, ranging from global warming to ozone depleti

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问题     Humans are thought to be responsible for a large number of
environmental problems, ranging from global warming to ozone
depletion. That is not in doubt, however, is the devastating effect【S1】______
humans are having on the animals and plants of the planet.
    Currently, an estimated 50,000 species become extinct each
year. If this takes on, the impact on all living creatures is likely to【S2】______
be profound. All species depend in some way each other to survive.【S3】______
If you remove one species from this very complex web of
interrelationships, you have very little idea about the knock-on
effects on the ecosystem. However, if you lose a key species, you【S4】______
might cause a whole flood of other extinctions.
    The most complicating matter is the fact in that there is no【S5】______
obvious solution to the problem. Unlike global warming and ozone
depletion—which, if the political will is there, could be reduced by【S6】______
cutting gas emissions—preserving bio-diversity remains an
intractable problem.
    The late idea is " sustainable management" , which involves【S7】______
using any species for human beings’ benefit, provide enough【S8】______
individuals of that species are left alive to ensure its continued
existence. Sustainable management is seen as a practical and
economic way of protecting species from extinction. Instead of【S9】______
depending on largely ineffective laws against illegal hunting, it gives
local people a good economic reason to preserve plants and animals.
    This sounds like a sensible strategy, but it remains to be seen
whether it will work. With corruption popular in many developing
countries, some observers are suspicious that the money will
actually reach people it is intended for. Others wonder how effective【S10】______
the locals will be at stopping illegal hunters.
【S1】

选项

答案That—What

解析 从句引导词误用。本句是含有主语从句和定语从句的复杂句,主语从句What is not in doubt中,what作该从句的主语,但that在引导主语从句时不在句子中作任何成分。
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