That large animals require a luxuriant vegetation, has been a general assumption which has passed from one work to another; but

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问题     That large animals require a luxuriant vegetation, has been a
general assumption which has passed from one work to another; but I do
not hesitate to say that it is completely true, and that it has vitiated the 【S1】______
reasoning of geologists on some points of great interest in the ancient
history of the world. The prejudice has probably derived from India, and 【S2】______
the Indian islands, which troops of elephants, noble forests, and 【S3】______
impenetrable jungles, are associated together in every one’s mind. If,
therefore, we refer to any work of travels through the southern parts of 【S4】______
Africa, we shall find illusions in almost every page either to the desert 【S5】______
character of the country, nor to the numbers of large animals inhabiting 【S6】______
it. The same thing is rendered evident by the many engravings which
have been published of various parts of the interior.
    Dr. Andrew Smith, who has lately succeeded in passing the Tropic
of Capricorn, informs me that, taken into consideration the whole of the 【S7】______
southern part of Africa, there can be no doubt of its being a sterile
country. On the southern coasts there are some fine forests, but without 【S8】______
these exceptions, the traveler may pass for days together through open
plains, covered by a poor and scanty vegetation. Now, if we look to the
animals inhabiting this wide plains, we shall find their numbers 【S9】______
extraordinarily great, and their bulk immense. We must enumerate the
elephant, three species of rhinoceros, the giraffe, two zebras, two
gnus, and several antelopes even larger than these latter animals. It may
be supposed that even although the species are numerous, the individuals 【S10】______
of each kind are few.
【S4】

选项

答案therefore—however

解析 衔接副词误用。该句意为“但是,如果我们以非洲南部旅行游记为参考的话,我们就会发现几乎游记的每一页要么提到了这个地区的沙漠地貌,要么是栖息在那里的众多大型动物。”此句与前句有明显的转折语气,因此将therefore改为however。
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